From: Using pooled data to estimate variance components and breeding values for traits affected by social interactions
σ w 2
m
One family
1 n σ P D 2 + 2 n − 1 σ P DI + n − 1 2 σ P I 2 + n − 1 r σ A T 2 − r σ A T 2
o/n
Two families
4 n σ P D 2 + 2 n − 1 σ P DI + n − 1 2 σ P I 2 + n 2 − 1 r σ A T 2 − r σ A T 2
2o/n
n families
n σ P D 2 + 2 n − 1 σ P DI + n − 1 2 σ P I 2 − r σ A T 2
o