Fig. 2
From: A theoretical derivation of response to selection with and without controlled mating in honeybees
Illustration of the genetic contributions to the breeding colonies with controlled mating. This figure motivates the recursion formula Eq. 5. Breeding colonies in year t receive half of their genetic material from their queens which as in the case of uncontrolled mating have an average breeding value equal to \(B_{t-2}+S_{1,t-2}\). The other half is inherited from the drones on a mating station whose common granddam on average has a breeding value equal to \(B_{t-3}+S_{2,t-3}\)