From: Comparison of linkage disequilibrium estimated from genotypes versus haplotypes for crossbred populations
Alleles at locus M
Alleles at locus N
0
1
Marginal frequency
\(r\)
\(s\)
\(r+s\)
\(t\)
\(u\)
\(t+u\)
\(r+t\)
\(s+u\)
\(r+s+t+u=1\)